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Publications

This is where you can read all of our latest publications including scientific studies, reports and research on plants and fungi.

Sarah Shuttleworth at DNA barcoding course
A waxcap mushroom growing in the grass, with mountains in the background

Discussion Paper on Ancient Grasslands

The Ancient grasslands report aims to create a practical definition of the term ‘Ancient Grassland’ to better identify and protect these important grassland habitats. It suggests multiple ways of defining the term and discusses possible advantages and disadvantages in using such definitions.

Definitions are suggested based on age of grassland as well as taking into consideration ecological, environmental and historical factors and landscape features. The report examines how the terminology we use with grasslands could aid their conservation in future.

Authors: A British Ecological Society report. Facilitated by Daniela Russi, Sarah Mckain, Rob Booth and Rebecca Walley. Contributions by Caroline O’Rourke, Professor David Gowing, Professor Gareth Griffith and Dr Kevin Walker. (Full list in annex).

Read the full report here

Looking out across the valley with Meadow Saxifrage in the foreground

Review of Trends in Grasslands Across the UK

This report explores the definition of grassland types across the UK and looks in to how these grasslands have changed in status and extent over the past century. Differing definitions can lead to difficulties in determining conservation status of grassland habitats. This report looks to explore the value of establishing consistent definitions of grassland types to aid in conservation efforts. 

The report also explores the commonly quoted statistic “around 97% of species-rich grasslands have been lost in the UK over the last 100 years” to look in to any supporting evidence. It investigates pressures on grasslands and reasons behind declines as well as exploring instances where grassland types have increased in area. 

Authors: Report by SLR consulting. Report commissioned by Plantlife and WWF-UK. 

Read the full report here

Valuing the Vital Grassland Ecosystem Services in the UK

This report offers a review of existing literature and evidence on the numerous advantages associated with species-rich grasslands. It focuses on the ecosystem services grasslands provide and their ability to contribute towards human wellbeing. 

This report examines how restoration and creation of semi-natural grasslands can offer opportunities to enhance ecosystem services while also providing a way of making these landscapes more resilient. It investigates and attempts to demonstrate the connection between grassland condition and future ecosystem services delivery. It points out how the services grasslands provide are often ‘invisible’ and as a result, grasslands can be overlooked and undervalued. 

Authors: Report by SLR consulting. Commissioned by Plantlife and WWF-UK. 

Read the full report here

A Bryophyte Red Data List for Wales

Wales supports almost 3 quarters of all British bryophyte species, therefore, there is a special responsibility to care for them. This report assesses threats to Welsh bryophytes, including mosses, liverworts and hornworts. It explores how they play a critical role in Welsh ecosystems, especially regarding water regulation and nutrient cycling.

This follows the publication of ‘A Vascular Plant Red Data List for Wales (Dines, 2008)’ to improve knowledge of the threats facing these species and to improve the availability of data. It’s the first report to assign threat categories to Welsh bryophytes and the first to use IUCN categories at a regional scale for this taxon group in Great Britain. The report discusses how IUCN categories were assigned. It also presents and explains the Red Data List for Wales.

Authors: Sam Bosanquet, Countryside Council for Wales, British Bryological Society, Trevor Dines, PLantlife Cymru.

Read the full report here

A Limestone Fern

A Vascular Plant Red Data List for Wales

This report assesses threats to Welsh vascular plants.

It’s the first report to assess the Welsh vascular plant flora using international criteria. The results of this report have been used to inform the selection of priorities for conservation in Wales and the report also allows a comparison between species at Great Britain and country level.

The report discusses how IUCN categories were assigned and then analyses differences between the Welsh list and the GB list.

The red data list for Wales is included at the end of this report and updates will be provided electronically as new data becomes available.

Authors: Dr Trevor Dines, Plantlife Wales. With help from the Vice-county recorders and Committee of the Botanical Society of the British Isles in Wales and the Countryside Council for Wales.

Read the full report here

Beautifully detailed photograph showing a purple coloured mushroom poking up from the forest floor

Important Fungus Areas

A provisional assessment of the best places for fungi in the UK.  

Fungi are of fundamental ecological importance and yet their conservation has been halted by a lack of site-based information. This report summarises a project which involved nominating sites to be considered as Important Fungus Areas (IFAs). The project resulted in a list of 520 sites which will be used to support and underpin biodiversity conservation in the UK. 

The report discusses some of the difficulties faced when surveying fungi and highlights the neglect that fungi have received with regards to their lack of inclusion in endangered species lists. 

It discusses the concept behind Important Fungus Areas and examines data collection methods and the criteria for selecting such areas. The full results are presented in the annex to this report. 

Authors: Report researched and written by Shelley Evans, Peter Marren and Martin Harper. 

Read the full report here

See more of our work here

Everything You Need to Know About Fungi

Fungi are often called the forgotten kingdom. Neither plants nor animals, fungi is a kingdom all of it’s own – and it’s a very important one indeed!

  • Go to:

We are on a mission to protect our precious fungi and the habitats they call home, and raise their profile by sharing this incredible hidden world. From insightful articles to tops tips as well ID guides and resources, this is where you’ll discover everything you need, to find out more about our fascinating fungi.

Read on below to learn about our current research projects, aiming to unlock more of this wonderful world. And find out how you can get involved helping us learn more about the fungi kingdom, by taking part in surveys as citizen scientists.

Our specialist botanical advisor Sarah Shuttleworth revisits her favourite fungi find – the rare Blue Tooth fungus. Sarah discusses what happened after her discovery and how, this and other fungi finds, can help to form a more detailed picture of an area, it’s history and inform how we can better protect these habitats for the future.

How to help protect fungi

Take Part in Waxcap Watch
Pink waxcap fungi growing in short green lawn

Take Part in Waxcap Watch

Britain is home to some of the most important waxcap grasslands in the world. However many species are becoming rare and declining; they need identifying and protecting. Become a citizen scientist and help us survey sites looking for waxcaps. You don’t need any fungi ID skills for a waxcap survey – all you need is a smart phone or access to a computer!

How to Find and Identify Waxcaps

How to Find and Identify Waxcaps

Get our top tips for finding grassland fungi, from where to look to what they look like! Use this guide to learn more about our wonderful waxcaps and help to get you on the road to finding, IDing and maybe even helping us to survey these incredible species.

Protecting Waxcaps: All the Losses We Cannot See…
A red coloured waxcap mushroom among grass

Protecting Waxcaps: All the Losses We Cannot See…

Discover the pressures the colourful waxcaps and their habitats face, the impact of development and what happens when there is a lack of data on the special sites these species call home. Here we also delve into how you can take action to protect them for the future.

Learn more about fungi

How Fungi Find Love on the Wood Wide Web

How Fungi Find Love on the Wood Wide Web

Join Senior Ecologist Sarah Shuttleworth for a deadwood date, as she shares what gets fungi swiping right on the wood wide web. Learn more about the connection that plants and fungi share.

Finding Hazel Gloves Fungus: Why Recording Matters

Finding Hazel Gloves Fungus: Why Recording Matters

Hazel Gloves Fungus is a priority species on the UK Biodiversity Action Plan, and a rare find for any fungi fan. Sarah Shuttleworth discovers this funky fungi for Reverse the Red month, and the secrets it reveals about the area it’s found in.

Waxcap Training Course
A pink mushroom growing in grass

Waxcap Training Course

Take your waxcap knowledge to the next level with our training course, suitable for keen fungi finders and land owners.

ID Guides

Spotter Sheets

Discover Fungi Species

Honey Fungus (Armillaria species) 
Honey fungus, Armillaria mellea, photographed on a dead branch in a woodland. The pale creamy brown of the fungus stands out against bright green mosses that cover the branch

Honey Fungus (Armillaria species) 

Ten wild, wacky and wonderful facts about the world of fungi

Here we delve into this mind-boggling realm to discover ten of weirdest, wackiest and most wonderful facts from the world of fungi! 

Photo showing a Yellow Brain fungus on a piece of dead wood on the forest floor. The bright yellow of the fungus stands out against the dark wood and leaf covered floor.

There at least five kingdoms in our living world, including; animals, plants, fungi, protists, and the prokaryotes (which includes bacteria). While the animal and plant kingdoms are well known, fungi are often overlooked – but the world of fungi is fascinating! And they’re all around us. A network of fungi runs through the earth in every continent and can even be found in our seas.  

Here we explore more of this incredible kingdom to share our passion for all things fungi. From a distant ancestral connection to how some fungi can survive on radioactive sites – read on for 10 wild, wacky and wonderful facts about the world of fungi!

1. The largest organism in the world is a fungus

Known as the Humungous Fungus, a single Armillaria organism, found in Oregon, is approximately 931 square hectares or around 70 football pitches! 

The fungus is measured including its network of mycelium that run underground, so much of the mass cannot be seen. While it’s unlikely you’ll come across anything of quite that size, there are some pretty big fungi you might find on your doorstep.

An illustration of a cream coloured Giant Puffball sat on a grass moundFor instance, another fungus species, the Giant Puffball Calvatia gigantea (shown in the illustration below) produces large, round fruiting bodies that can be seen above ground. They’re certainly hard to miss, as they can grow up to 80cms in diameter (outside of the UK, they have been known to grow as big as 1.5m). They can grow to such a size that on occasion, people have mistaken them for sheep in a field!

2. Fungi and humans share a distant ancestor

Fungi is a kingdom all of its own, so we don’t want to mistake it for being a plant or animal. That said, while you might expect that these interesting organisms are more closely related to plants – the opposite is true. Fungi share a common ancestor with the animal kingdom, which includes humans, dating back around 1 billion years. [1]

While that may seem an incomprehensible length of time, it might not be in the world of fungi. We don’t have conclusive evidence of how old fungi can be. Research suggests that some of the fungi on our planet today, have been alive for thousands of years. 

The ‘Humungous Fungus’ is estimated to be between 1,900 and 8,650 years old! [2] Which means there could be living fungi that were present when the Vikings landed in the UK, when the Romans ruled and even when Stonehenge was erected. 

3. Fungi can be found all around your kitchen 

Fungi might be a common feature on your breakfast plate, or in a bowl of pasta – but there are many other foods that contain fungi, which might surprise you. 

Did you know that bread, beer, wine and some cheeses all include types of fungi? Yeast-forming fungi are an essential ingredient in the processes to make them and there are many different species, the most common of which is Saccharomyces cerevisiae, used in brewing and making bread.  

As far as we know today there are more than 2,000 species of edible fungi in the world. [3] These range from the popular Button and Chestnut Mushrooms Agaricus bisporus to the mycoprotein that makes up the meat substitute Quorn. And they are becoming even more popular with Lion’s Mane Hericium erinaceus being added to some coffees and fungi being used to help ferment cacao beans to make chocolate!  

Photo shows a number of clear test tubes resting in a yellow tray. There is a plant in a vase behind the test tubes.

4. Fungi are friends not (just) food

That’s right, fungi aren’t just contained to your kitchen, you can probably find them all around house from your wardrobe to your medicine cabinet and even your beauty bag. 

One common example of fungi used in medicine is Penicillin, the first antibiotic, which was derived from Penicillium mould. Other fungi produce compounds that are used to treat diseases like cancer, high cholesterol and even organ transplant rejection. 

In the beauty industry mushrooms are becoming a common feature of skincare products as many have anti-inflammatory and antioxidant properties. They have even been trialled as a renewable replacement for dyes in cosmetics. 

Fungal enzymes are also used in the processing and production of a variety of products including paper, leather, cotton and biofuels – and generally they replace harsher chemicals in these processes. 

5. Fungi are helping our planet – in more ways than one

Fungi are essential for the nutrient cycles of the planet by acting as natural decomposers for organic matter. This means that as plants die, fungi help to break them down. In fact fungi are the only group of organisms that can break down plant cell walls. 

But research suggests that fungi could also break down other substances, in a process which could be used to clean our planet. Mycoremediation, is a process that uses fungi to break down substances that cause pollution. 

This process can be seen in action in places like the Amazon rainforest where local landowners are using these techniques to rid the ground of crude oil contamination. 

Currently researchers are also using fungi to create filters to clean water. They’ve even found a species called Aspergillus tubingensis, which can break down plastics! [4]

And five years after the nuclear disaster at Chernobyl, the fungi Cryptococcus neoformans, was found feeding on radiation. [5] Scientists are now exploring the possibilities of this incredible fungus, looking at whether its abilities could help protect astronauts from radiation in space. 

Photo shows lots of of Sulphur Tuft mushrooms on the bottom of a tree. The photo is taken in the day and shows the beige or tan colouring of the small umbrella-shaped mushrooms.

6. There are fungi that can glow in the dark

Bioluminescence is a phenomenon where a living organism is able to produce and emit light. You might have seen this trait in fireflies in the sky or algae in the water, but more than 70 types of fungi can also glow in the dark. [6]

Some of these fascinating fungi that can be found around the world include Sulphur Tuft Hypholoma fasciculare (pictured on the right, in the daytime), Jack-O’Lantern Mushroom Omphalotus olearius, and Little Ping-Pong Bats Panellus pusillus. 

You can even find some of these luminescent fungi in the UK – though they are rarer and weaker here than in other parts of the world. 

Photograph shows a fallen, decaying tree of the forest floor, covered in hundreds of mushrooms.

7. Fungi store carbon

While plants have long been heralded for their ability to store carbon, they are not the only organisms capable of doing so. 

Fungi help break down organic material, using the carbon recycled by plants, and store this carbon as mycelium underground. This process helps lock atmospheric carbon in the soil, as well as releasing other nutrients into the soil which can help plants grow.  

Scientists estimate that fungi could be storing up to 13.12 billion metric tons of carbon annually! [7]

8. People have been using fungi for thousands of years

We might still be finding new and exciting ways to benefit from fungi today, but our ancestors paved the way.  

Most notably, a number of harvested fungi were found in the bag of Otzi ‘The Iceman’. Otzi’s mummified remains were discovered on the Italian-Austrian border in 1991. He had different species of fungi with him, though the intended uses of these cannot be known. Otzi is estimated to have been alive between 3350 and 3105 BC, meaning humans have been connected to fungi for at least 5,000 years!

Photograph shows a few Jelly Ear Fungi on tree, that is also covered in moss. The fungi are a dark brown in colour and almost perfectly resemble the shape of a human ear.

9. Fungi come in all shapes and sizes

While we tend to think of fungi in that stereotypical umbrella form, they can come in a variety of different sizes and shapes – including some pretty impressive lookalikes! 

Take the Jelly Ear Fungus Auricularia auricula – a common fungus that you can often find on decaying or dead Elder. This odd fungus gets its name because it looks and feels, almost completely like a human ear! 

Then there’s Dead Man’s Fingers Xylaria polymorpha! Just about as creepy visually as it sounds. The fruiting bodies are long and club shaped, and could easily be mistaken for a hand poking up out of the soil. 

Finally, a much less scary lookalike, is the Bird’s Nest Fungus Crucibulum laeve – named because it’s fruiting body almost perfectly resembles a tiny bird’s nest filled with eggs! When it rains, drops splash into the nest shape and knock out one of the little discs, which contain the spores, helping them to spread. 

10. Fungi might be able to make their own ‘weather’

OK, it’s all relative – you’re not going to find a mushroom that brings out the blue skies or creates a thunderstorm over your town. But there is research to suggest that some species, including Shitake Lentinula edodes and Oyster Mushrooms Pleurotus ostreatus could create their own mini weather systems. [8]

The science behind this phenomenon is evaporative cooling. Mushrooms release water, which evaporates in the warm air, turning into water vapour. The water vapour then cools the air, which sinks and the warm air in turn rises. As the air moves it creates a small breeze, probably not one that we would notice – but it’s enough for the fungi. 

Even a whisper of a breeze can help to disperse the tiny spores and spread them over a larger distance. This trick is especially helpful on hot days where there isn’t any wind to assist.

  • References

    1. Read more about animals distant connection to fungi in this report. 
    2. Read more about the ‘Humongous Fungus’ in this report. 
    3. Read more about edible fungi in this report. 
    4. Read more about how fungi can help clean water and break down plastics here. 
    5. Read about how fungi have been surviving in radiation here. 
    6. Learn about glowing fungi here.  
    7. Read more about how fungi store carbon here. 
    8. Read more about fungi’s own weather systems here. 

Read more

Rare Fungus spotted at Kenfig National Nature Reserve
Brown topped fungus with yellow gills in a green grassy area.

Rare Fungus spotted at Kenfig National Nature Reserve

Chris Jones, the Warden of Kenfig National Nature Reserve, recently found the very rare fungus, during a routine survey.

Half-term Idea: Family Fungi Fun
Girl looking at fungi

Half-term Idea: Family Fungi Fun

Fungi are one of our most fascinating creations, and best of all, they are right under our noses – perfect for children to spot.

Protecting Waxcaps: All the Losses We Cannot See…

Protecting Waxcaps: All the Losses We Cannot See…

Britain’s waxcap grasslands are considered to be the best in Europe. Discover the pressures these colourful fungi and their habitats face…

Which seed should I use?

Whether you wish to create a meadow from scratch or introduce more wildflowers and grasses into your meadow or grassy area, or within your garden lawn, our species lists are here to guide you.

A wildflower meadow with yellow, white, purple flowers in among the grass. The meadow is in Ryewater, Dorset. Image is by Jo Costley.

When selecting seeds, the first things to think about are:

  • soil type
  • fertility levels
  • problem plants and how to manage them.

You can do this by understanding what the species that are already there are telling you about your soil, fertility and its existing species, and/or by carrying out a soil test. If you are wishing to introduce more species to an existing grassland, then knowing what you already have is important so that you can tailor what you introduce to be an appropriate match.

It is best to survey your grassy area over the summer and use our handy ‘plant forensics’ guide to help you understand which species are likely to do well, before purchasing any seed.

We recommend using local seed mixes which you can check against the most appropriate species list below to be sure of a good match. Alternatively, if you know a local meadow with appropriate species, and you can organise either green hay or brush-harvesting, or even hand collecting ripe seed of specific species, this would be even better.

Choosing your species

field full of variety of flowers in pink, yellow purple

To help you on your meadow-making journey, we have made these lists as a starting point for creating or restoring grassland in a typical species-poor or species-moderate grassland scenario.

Please note that these are not definitive lists, and different sites and situations may require a more bespoke approach. If your grassland is outside of the ‘norm’ such as within a national park or AONB – then contact us for a more bespoke guidance on seed-mixes which will reflect the site’s regional distinctiveness.

For example, a pH neutral hay meadow in the south west of England won’t have the same community as a neutral hay meadow in the Lake District . We have also left out species which are very geographically specific. So, if you have an unusual site with potential to support a rare habitat, then contact us for more bespoke list.

These lists divide the species into Groups 1, 2 and 3, in order of both their difficulty establishing from seed-introduction, and their tolerance to high soil-nutrient levels.

  • Group 1 – tolerant of higher nutrient levels and easy to establish from seed
  • Group 2 – tolerant of moderate nutrient levels, moderate colonisers from seed
  • Group 3 – poor colonisers, sensitive and poor soils only, may need introducing via established plugs

No quantities have been outlined in the lists below because seed providers should be able to provide this detail, or already have this outlined in their meadow-mixes. In general, however, grassland communities will be between 50-80% grass, and those with high nutrient levels will usually settle into a higher percentage content of the grass species.

Species lists for different grassland types

pH Neutral grassland 5-6.5

  • All these grasses can be introduced as seed

    Grasses to introduce (if absent):

      • Yellow Oat-grass – Trisetum flavescens
      • Smooth Meadow-grassPoa pratensis 
      • Meadow FoxtailAlopecurus pratensis 
      • Smaller Cat’s-tailPhleum bertolonii 
    Grasses (additional) if bare soil:

      • Yorkshire-fogHolcus lanatus 
      • Red FescueFestuca rubra 
      • Crested Dog’s-tailCynoscurus cristatus 
      • Common BentAgrostis capillaris 
      • Sweet Vernal-grassAnthoxanthum odoratum 
      • Meadow BarleyHordeum secalinum 
  • Group 1

    • Black/Common KnapweedCentaurea nigra 
    • Burnet SaxifragePimpinella saxifrage 
    • Common Birds-foot TrefoilLotus corniculatus 
    • Meadow Vetchling – Lathyrus pratensis 
    • Ox-eye DaisyLeucanthemum vulgare 
    • Rough HawkbitLeontodon hispidus 
    • Bulbous ButtercupRanunculus bulbosus 
    • Common Cat’s-earHypochaeris radicata 
    • Common SorrelRumex acetosa 
    • Lesser TrefoilTrifolium dubium 
    • Ribwort PlantainPlantago lanceolata 
    • Meadow ButtercupRanunculus acris 
    • Red Clover (native variety)Trifolium pratense var pratense 
    • SelfhealPrunella vulgaris
    • YarrowAchillea millifolium 
    • Wild Carrot – Daucus carota 
    • Germander SpeedwellVeronica chamaedrys 
    • Black MedickMedicago lupilina 

     

  • Group 2

    • Autumn HawkbitScorzoneroides autumnalis 
    • Goat’s-beardTragopogon pratensis 
    • Musk MallowMalva moschata 
    • AgrimonyAgrimony eupatoria 
    • CowslipPrimula veris 
    • Lady’s BedstrawGallium verum 
    • Salad BurnetPoterium sanguisorba 
    • Yellow RattleRhinanthus minor 
    • BugleAjuga reptans 
    • Tufted Vetch – Viccia cracca 
    • Eyebright sp – Euphrasia sp. 
    • Common Milkwort – Polygala vulgaris 
    • TormentilPotentilla erecta 
    • PignutConopodium majus 

     

  • Group 3

    • Betony – Stachys officinalis
    • Common Spotted-orchidDactylohiza fuchsii (If more alkaline)
    • Bee Orchid – Ophrys apifera 
    • Pyramidal OrchidAnacamptis pyramidalis (If more alkaline)
    • Dyer’s GreenweedGenista tinctoria 
    • Wood AnemoneAnemone nemorosa 
    • Field ScabiousKnautia arvensis 

     

Wetter soils/floodplain (neutral pH 5-6.5)

  • All these grasses can be introduced as seed

    Grasses to introduce (if not present):

    • Meadow FoxtailAlopecurus pratensis 
    • Sweet Vernal-grass  – Anthoxanthum odoratum 
    • Marsh FoxtailAlopecurus geniculatus 
    • Crested Dog’s-tailCynosurus cristatus 
    Grasses (additional) if bare soil:

    • Creeping BentAgrostis stolonifera 
    • Yorkshire-fogHolcus lanatus 
    • Red FescueFestuca rubra agg. 
  • Group 1

    • Ragged RobinSilene flos-cuculi 
    • CuckooflowerCardamine pratensis 
    • Great BurnetSanguisorba officinalis 
    • Greater Bird’s-foot TrefoilLotus pedunculatus 
    • Common SorrelRumex acetosa 
    • Lesser TrefoilTrifolium dubium 
    • AngelicaAngelica sylvestris 
    • Ribwort PlantainPlantago lanceolata 
    • Meadow ButtercupRanunculus acris 
    • Red Clover (native variety)Trifolium pratense var pratense 
    • Common FleabanePulicaria dysenterica 

     

  • Group 2

    • MeadowsweetFilipendula ulmaria 
    • Marsh/Fen BedstrawGallium uliginosum 
    • Water MintMentha aquatica 
    • Common BistortBistorta officinalis 
    • Common Meadow-rueThalictrum flavum 
    • Pepper SaxifrageSilaum silaus 
    • SneezewortAchillea ptarmica 
    • Creeping JennyLysimachia nummularia
    • Water AvensGeum rivale 
    • Narrow-leaved Water-dropwortOenanthe silaifolia 
    • Tufted VetchViccia cracca 
    • Marsh MarigoldCaltha palustris 

     

  • Group 3

    • Devil’s-bit ScabiousSuccisa pratensis 
    • Saw-wortSerratula tinctorium 
    • BetonyStachys officinalis 
    • Marsh SpeedwellVeronica scutellata 
    • Marsh ValerianValeriana dioica 
    • Southern Marsh OrchidDactylorhiza praetermissa (where more alkaline)

     

Lowland acidic grassland (pH < 5.5)

  • All these grasses can be introduced as seed

    Grasses to introduce (if absent):

    • Sweet Vernal-grassAnthoxanthum odoratum 
    • Common BentAgrostis capillaris 
    • Red FescueFestuca rubra 
    • Sheep’s FescueFestuca ovina 
    • Wavy Hair-grassDeschampsia flexuosa 
    • Squirrel-tail FescueVulpia bromoides 
     

    • Early Hair-grassAira praecox 
    • Silver Hair-grassAira caryophyllea 
    • Heath GrassDanthonia decumbens 
    • Fine-leaved Sheep’s fescueFestuca filiformis 
    • Velvet Bent (where damp)Agrostis canina  
    • Brown BentAgrostis vinealis 
  • Group 1

    • Common Bird’s-foot Trefoil – Lotus corniculatus 
    • Rough HawkbitLeontodon hispidus 
    • Field WoodrushLuzula campestris 
    Where mildly acidic:

    • Common Cat’s-earHypochaeris radicata 
    • Lesser TrefoilTrifolium dubium 
    • Ribwort PlantainPlantago lanceolata 
    • SelfhealPrunella vulgaris 
    • YarrowAchillea millifolium 
    • Black MedickMedicago lupilina 

     

  • Group 2

    • Sheep’s SorrelRumex acetosella 
    • Heather/Ling  – Calluna vulgaris 
    • Heath BedstrawGallium saxatile 
    • Lady’s Bedstraw (on mild acid)Gallium verum 
    • Lesser StitchwortStellaria graminea 
    • Bell HeatherErica cinerea
    • Common Bird’s-footOrnithopus perpusillus 
    • Common CentauryCentaurium erythraea 
    • Common Stork’s-billErodium cicutarium 
    • Heath MilkwortPolygala serpyllfolia 
    • Mouse-ear HawkweedPilosella officinarum 
    • TormentilPotentilla erecta 
    • Common Dog-violetViola reichenbechiana 
    • Heath VioletViola canina 
    • Autumn HawkbitScorzoneroides autumnalis (where mildly acidic)
    • PignutConopodium majus 

     

  • Group 3

    • Betony – Betonica officinalis
    • Devil’s-bit Scabious Succisa pratensis
    • Heath Spotted-orchidDactylorhiza maculata 
    • Wild ThymeThymus polytrichus 
    • Blue FleabaneErigeron acer 
    • Wood AnemoneAnemone nemorosa 
    • HarebellCampanula rotundifolia 
    • Heath Speedwell (on anthills)Veronica officinalis 
    • Field ScabiousKnautia arvensis 
    • Bitter-vetchLathyrus linifolius 
    • Lesser HawkbitLeontodon saxatilis 
    • Biting StonecropSedum acre 
    • LousewortPedicularis sylvatica 
    • Saw-wortSerratula tinctoria 
    • Wood SageTeucrium scorodonia 
    • Sheep’s-bitJasione montana 

     

Lowland Calcareous pH > 6.5

  • All these grasses can be introduced as seed

    Grasses to introduce (if absent):

    • Upright Brome – Bromopsis erecta 
    • Crested Dog’s-tailCynosurus cristatus 
    • Red FescueFestuca rubra agg. 
    • Downy Oat-grassHelictotrichon pubescens 
    • Meadow Oat-grassHelictotrichon pratense 
    • Yellow Oat-grassTrisetum flavescens
    • Narrow-leaved Meadow-grassPoa angustifolia 
    • Quaking-grassBriza media 
    • Crested Hair-grassKoeleria macrantha 
    • Yellow Oat-grassTrisetum flavescens 
    • Sheep’s FescueFestuca ovina 
  • Group 1

    • Common Bird’s-foot Trefoil – Lotus corniculatus 
    • Oxeye DaisyLeucanthemum vulgare 
    • Rough HawkbitLeontodon hispidus
    • Kidney VetchAnthyllis vulneraria 
    • Chalk KnapweedCentaurea debeauxii 

    Where mildly alkaline:

    • Bulbous ButtercupRanunculus bulbosus 
    • Common Cat’s-earHypochaeris radicata 
    • Common SorrelRumex acetosa 
    • Lesser TrefoilTrifolium dubium 
    • Ribwort PlantainPlantago lanceolata 
    • Red Clover (native variety)Trifolium pratense var pratense 
    • SelfhealPrunella vulgaris 
    • YarrowAchillea millifolium 
    • Germander SpeedwellVeronica chamaedrys
    • Black MedickMedicago lupilina 

     

  • Group 2

    • CowslipPrimula veris
    • Autumn HawkbitScorzoneroides autumnalis (where mildly alkaline)
    • Tufted VetchViccia cracca (where mildly alkaline)
    • Dwarf Thistle – Cirsium acaule 
    • Lady’s BedstrawGallium verum 
    • Sainfoin – Onobrychis viciifolia 
    • Salad BurnetPoterium sanguisorba 
    • Carline Thistle (s-facing)Carlina vulgaris  
    • Yellow RattleRhinanthus minor 
    • Common Centaury (well-drained)Centaurium erythraea 
    • Bladder Campion  – Silene vulgaris 
    • Common Stork’s-billErodium cicutarium 
    • Mouse-ear HawkweedPilosella officinarum 
    • Thyme-leaved SandwortArenaria serpyllifolia 
    • Wild Basil (southern UK)Clinopodium vulgare 
    • MarjoramOriganum vulgare 
    • Yellow-wortBlackstonia perfoliate 

     

  • Group 2 or 3

    • EyebrightsEuphrasia officinalis agg. 
    • Autumn GentianGentianella amarella 
    • English Gentian (only in south and midlands)Gentianella anglica 
    • Common MilkwortPolygala vulgaris 
    • Saw-wortSerratula tinctorium 
    • SquinancywortAsperula cynanchica 
    • Lesser HawkbitLeontodon saxatilis
    • Common Restharrow 
    • Wild Strawberry 

     

  • Group 3

    • Field ScabiousKnautia arvensis  
    • BetonyStachys officinalis
    • Clustered Bellflower 
    • Common Rock-roseHelianthemum nummularium 
    • Devil’s-bit ScabiousSuccisa pratensis 
    • Common Spotted-orchidDactylohiza fuchsii 
    • Pyramidal OrchidAnacamptis pyramidalis
    • Bee Orchid Ophrys apifera 
    • DropwortFilipendula vulgaris 
    • Fairy FlaxLinum catharticum 
    • Greater Knapweed 
    • Hairy Violet 
    • HarebellCampanula rotundifolia 
    • Hoary PlantainPlantago 
    • Horseshoe Vetch 
    • Small ScabiousScabiosa columbaria 
    • Wild ThymeThymus polytrichus 
    • Large ThymeThymus pulegioides 
    • Biting StonecropSedum acre 
    • Bloody Crane’s-bill 
    • Common MilkwortPolygala vulgaris 
    • Chalk Milkwort 

     

Upland habitats – all those grasslands which are above approximately 250m (sea level). If close to 250m and somewhat sheltered, the above lowland species may also thrive.

Upland calcareous pH > 6.5

  • All these grasses can be introduced as seed

    Grasses to introduce (if absent):

    • Upright Brome – Bromopsis erecta 
    • Crested Dog’s-tailCynosurus cristatus 
    • Red FescueFestuca rubra agg. 
    • Downy Oat-grassHelictotrichon pubescens 
    • Meadow Oat-grassHelictotrichon pratense 
    • Yellow Oat-grassTrisetum flavescens
    • Narrow-leaved Meadow-grassPoa angustifolia 
    • Quaking-grassBriza media 
    • Crested Hair-grassKoeleria macrantha 
    • Sheep’s FescueFestuca ovina 
  • Group 1

    • Common Bird’s-foot Trefoil – Lotus corniculatus 
    • Rough HawkbitLeontodon hispidus

     

  • Group 2

     

    • Salad BurnetPoterium sanguisorba 
    • Carline Thistle (s-facing)Carlina vulgaris 
    • Mouse-ear HawkweedPilosella officinarum 

     

  • Group 2 or 3

    • EyebrightsEuphrasia officinalis agg. 
    • Autumn GentianGentianella amarella 
    • SquinancywortAsperula cynanchica 

     

  • Group 3

    • Common Rock-roseHelianthemum nummularium 
    • DropwortFilipendula vulgaris 
    • Devil’s-bit ScabiousSuccisa pratensis 
    • Fairy FlaxLinum catharticum 
     

    • HarebellCampanula rotundifolia 
    • Wild ThymeThymus polytrichus 
    • Small Scabious Scabiosa columbaria 
    • Horseshoe VetchHippocrepis comosa 

     

Upland acid pH < 5.5

  • All these grasses can be introduced as seed

    Grasses to introduce (if absent):

    • Sweet Vernal-grassAnthoxanthum odoratum 
    • Common BentAgrostis capillaris 
    • Sheep’s FescueFestuca ovina 
    • Wavy Hair-grassDeschampsia flexuosa 
    • Yorkshire-fogHolcus lanatus 
    • Fine-leaved Sheep’s Fescue  – Festuca filiformis  
    • Velvet Bent (where damp)Agrostis canina 
    • Brown BentAgrostis vinealis 
    • Squirrel-tail FescueVulpia bromoides 
    • Early HairgrassAira praecox 
    • Silver Hair-grassAira caryophyllea 
    • Heath Grass Danthonia decumbens 
  • Group 1

    • Common Bird’s-foot Trefoil – Lotus corniculatus 
    • Field Woodrush – Luzula campestris 
    • Rough HawkbitLeontodon hispidus

     

  • Group 2

     

    • Sheep’s Sorrel – Rumex acetosella 
    • Heath BedstrawGallium saxatile 
    • Heather/LingCalluna vulgaris 
    • TormentilPotentilla erecta (group 2 or 3)

     

Upland hay meadows pH neutral 5 – 6.5

  • All these grasses can be introduced as seed

    Grasses to introduce (if absent):

     

    • Yellow Oat-grassTrisetum flavescens
    • Smooth Meadow-grassPoa pratensis 
    • Yorkshire-fogHolcus lanatus 
    • Crested Dog’s-tailCynosurus cristatus 
    • Red FescueFestuca rubra agg. 
     

    • Common Bent – Agrostis capillaris 
    • Sweet Vernal-grassAnthoxanthum odoratum 
    • Meadow BarleyHordeum secalinum 
  • Group 1

    • Common Bird’s-foot Trefoil – Lotus corniculatus 
    • Black/common KnapweedCentaurea nigra 
    • Burnet SaxifragePimpinella saxifrage 
    • Great Burnet Sanguisorba officinalis 
    • Rough HawkbitLeontodon hispidus
    • Ragged RobinSilene flos-cuculi 
    • Meadow VetchlingLathyrus pratensis 
    • Autumn HawkbitScorzoneroides autumnalis (group 1 or 2)

     

  • Group 2

     

    • BugleAjuga reptans 
    • MeadowsweetFilipendula ulmaria 
    • Lady’s MantlesAlchemilla sp. 
    • Marsh MarigoldCaltha palustris 
    • SneezewortAchillea ptarmica
    • Water AvensGeum rivale 
    • Common BistortBistorta officinalis 

     

     

  • Group 2 or 3

    • EyebrightsEuphrasia officinalis agg. 
    • TormentilPotentilla erecta
    • PignutConopodium majus 
    • Saw-wortSerratula tinctorium 
    • Yellow RattleRhinanthus minor 

     

  • Group 3

     

    • Devil’s-bit ScabiousSuccisa pratensis 
    • Wood Anemone Anemone nemorosa 
     

    • Wood Crane’s-bill 
    • Meloncholy ThistleCirsium heterophyllum  

     

More Meadow Guidance

  • Go to:

What’s a lichen?

Step 1.

Even though we can find lichens in most places we go, lots of us have never really noticed them and don’t know what they are.  

Have a go at our What’s a lichen? activity to learn everything you need to know about them. 

 

Have a go at our activity

Step 2.

Now you know what a lichen is, can you tell lichens apart from other living things?

Take a look at the activity sheet and see if you can Spot the lichen. 

 

Download and print Spot the lichen sheet

Scroll down for even more lichen activities!

Lichens are all around us!

Step 1.

Lichens can grow almost anywhere… they could even be right on your doorstep! 

Find out more about where lichens grow through our online activity – you might be surprised at where you can find them! 

 

Have a go at our activity

Step 2.

Even in busy places, full of people and cars, you’ll be able to spot lichens. They could be anywhere, from playparks and pavements, to graveyards and post boxes. 

Now you know some of the places you might find lichens growing its time to see if you can find some. Go on your own Lichen Hunt to see where you can find them in your local area.  

 

Download and print the Lichen Hunt sheet

Activity Sheets

How to spot it

A tall, strong smelling, hairy plant with a ridged stem. Oval toothed leaves with yellow umbrella shaped flowers.

Other Species

Publications
A person holding a camera about to take a photo of a plant.

Publications

This is where you can read all of our latest publications including scientific studies, reports and research.

Everything you need to know about fungi

Everything you need to know about fungi

We are on a mission to protect fungi, the habitats they call home and raise their profile across the world.

Managing Meadows
Wildflower meadow bursting with yellow dandelions and rolling hills in the background

Managing Meadows

Meadows are often a defining feature of summer, but when managed properly they can also be a vital habitat for wildlife and nature recovery.